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Rules of Nomenclature



Classification is more than just naming things. Classifications have a function.

Biosystematics alone among the sciences tends to stress diversity, rather than commonality among organisms. It is a far less REDUCTIONIST field than others in the sciences.


Levels of Taxonomy

Why should you care about biosystematics?

Because knowledge of classification at the alpha, beta and gamma levels can save you and the ecosystems around you.

Let's look at an example...

Another example:

  • 1920's - Hawaiian forest preserves' fern populations were severely threatened by the invasion of an exotic fern weevil (Syagrius fulvitarsis). No one knew where it was native, and it was highly reproductive and difficult to control.

  • 1921 - systematist Pemberton identified a single museum specimen, labeled with locality, as Australian.

  • The beetle's natural predators from Australia could then be determined and (after careful study!) used as biological control.


    Some important definitions for future use...


    Taxon has dimensions in space (geographical range) and time (its evolutionary history).

    Example we'll use: Family Pongidae, the Great Apes.

    To determine the symplesiomorphies that link this group together, we use an OUTGROUP, the next most recent relative of the entire assemblage above. In this case, an appropriate outgroup would be the Old World Monkeys.

    The more symplesiomorphies a taxon within your study group shares with the outgroup, the more likely it is that it has a recency of common descent with that outgroup, and may be more primitive with respect to the other members of the assemblage.

    Only SYNAPOMORPHIES can be used to establish recency of common descent among related organisms sharing many plesiomorphies.

    SYMPLESIOMORPHIES help us establish that a study group shares characters with a hypothetical ancestor, but only the SYNAPOMORPHIES exhibited by each group tell us about how closely related they are to each other.

    The more synapomorphies two groups exhibit, the more recent their common ancestor. For example, from the phylogenetic tree of the Great Apes we drew in class, you can surmise that humans and Bonobos exhibit more synapomorphies (shared, derived characters) than do humans and chimpanzees. And similarly, that Gorillas, chimps, Bonobos and humans exhibit more synapomorphies (as a group) than do Gibbons and humans.

    A taxon's evolutionary history/relationships can be diagrammed with a phylogenetic tree, similar to the one you constructed in your BIOSYSTEMATICS WORKSHOP.

    Form taxon: a taxon whose members are included in the group more on the basis of shared, known similarities in morphology, physiology, etc. than on known evolutionary relationships. (e.g., Kingdom Protista, Kingdom Monera, Division (Phylum) Deuteromycota)

    THREE SCHOOLS OF THOUGHT IN EVOLUTIONARY BIOLOGY

    (Be sure to review this in your Lab Manual)

  • Classical Evolutionary

  • Phenetic

  • Cladistic


    Classical Evolutionary Taxonomy

  • Was once the most commonly taught philosophy

  • both common ancestry and time of evolutionary diversification since splitting from a common ancestor are considered important.

  • specialization after a branch point is considered important

  • monophyletic & paraphyletic groups are acceptable

  • This method tends to be less objective than cladistic system; investigator bias has often clouded true evolutionary relationships


    The Phenetic System

  • monophyly, paraphyly and polyphyly have no meaning to the pheneticist (a.k.a. "numerical taxonomist"). The hard core pheneticist might assert that because true evolutionary relationships are not possible to know (we can't go back and make sure), that such terms are irrelevant and impossible to confirm.


    The Cladistic system

    Four tenets of cladism:


    To the Cladist: