Professor Emeritus

My current research centers around three major themes: evaluation of historical and ecologic determinants of the distributions of tropical organisms (especially amphibians and reptiles); their ecological role in tropical forests; and biogeographic theory.

My most recent publication is a comprehensive handbook on the 385 species of Costa Rican amphibians and reptiles. The book includes keys, diagnostic descriptions, ecological data and a biogeographic analysis of the herpetofauna. The well-known nature photographers Michael and Patricia Fogden provide the book with full color illustrations.



Awards and Recognitions:

  • 1974-1980 Founder(1963) and President of the Organization for Tropical Studies
  • 1982-2000 Member of the International Commission on Zoological Nomenclature
  • President of the Southern California Academy of Sciences
  • President of the American Society of Ichthyologists and Herpetologists
  • President of the Society of Systematic Biologists
  • 1998 Honorary Member of Academia Nacional de Ciencias de Costa Rica
  • 2000 Henry S. Fitch Award for Excellence in Herpetology
  • Archie F. Carr, Jr. Medal from the Florida Museum of Natural History and the University of Florida
  • 2005 Outstanding Service Award of the American Institute of Biological Sciences
  • 2009 Robert K. Johnson Distinguished Service Award of the American Society of Ichthyologists and Herpetologists
  • Recent Representative Publications:

    Send email to savy1@cox.net

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